Turtle


Turtles are an order of reptiles known as Testudines, characterized by a shell developed mainly from their ribs. Modern turtles are divided into two major groups, the side-necked turtles and hidden neck turtles which differ in the way the head retracts. There are 360 living and recently extinct species of turtles, including tortoises and terrapins. They are found on most continents, some islands and much of the ocean. Like other reptiles, birds, and mammals, they breathe air and do not lay eggs underwater, although many species live in or around water. Genetic evidence typically places them in close relation to crocodilians and birds.

Turtle shells are made mostly of bone; the upper part is the domed carapace, while the underside is the flatter plastron or belly-plate. Its outer surface is covered in scales made of keratin, the material of hair, horns, and claws. The carapace bones develop from ribs that grow sideways and develop into broad flat plates that join up to cover the body. Turtles are ectotherms or "cold-blooded", meaning that their internal temperature varies with their direct environment. They are generally opportunistic omnivores and mainly feed on plants and animals with limited movements. Many turtles migrate short distances seasonally. Sea turtles are the only reptiles that migrate long distances to lay their eggs on a favored beach.

Turtles have appeared in myths and folktales around the world. Some terrestrial and freshwater species are widely kept as pets. Turtles have been hunted for their meat, for use in traditional medicine, and for their shells. Sea turtles are often killed accidentally as bycatch in fishing nets. Turtle habitats around the world are being destroyed. As a result of these pressures, many species are threatened with extinction.

The word turtle is derived from the French tortue or tortre ('turtle, tortoise').[3] It is a common name and may be used without knowledge of taxonomic distinctions. In North America, it may denote the order as a whole. In Britain, the name is used for sea turtles as opposed to freshwater terrapins and heavy-footed, land-dwelling tortoises. In Australia, which lacks true tortoises (family Testudinidae), non-marine turtles were traditionally called tortoises, but more recently turtle has been used for the entire group.[4]

The name of the order, Testudines (/tɛˈstjdɪnz/), is based on the Latin word for tortoise, testudo;[5] and was coined by German naturalist August Batsch in 1788.[1] The order has also been historically known as Chelonii (Latreille 1800) and Chelonia (Ross and Macartney 1802)[2] which are based on the Ancient Greek word for tortoise: χελώνη (chelone).[6][7] Testudines is the official order name due to the Principle of Priority.[2] The term chelonian is used as a formal name for members of the group.[1][8]

The largest living species of turtle (and fourth-largest reptile) is the leatherback turtle which can reach over 2.7 m (8 ft 10 in) in length and weigh over 500 kg (1,100 lb).[9] The largest known turtle was Archelon ischyros, a Late Cretaceous sea turtle up to 4.5 m (15 ft) long, 5.25 m (17 ft) wide between the tips of the front flippers, and estimated to have weighed over 2,200 kg (4,900 lb).[10] The smallest living turtle is Chersobius signatus of South Africa, measuring no more than 10 cm (3.9 in) in length[11] and weighing 172 g (6.1 oz).[12]


Photograph of one half of a tortoise skeleton, cut in half vertically showing the vertebrae following curving along the carapace
Sagittal section of a tortoise skeleton
Development of the shell. The ribs are growing sideways into the carapacial ridge, seen here as a bud, to support the carapace.[16]
Head and neck of African helmeted turtle
Sea turtles have streamlined shells and limbs adapted for fast and efficient swimming.[31]
The red-eared slider has an exceptional seven types of color-detecting cells in its eyes.[34]
A submerged Indian softshell turtle nose-breathing at river surface
Snapping turtle emerging from period of brumation, in which it buried itself in mud. Turtles have multiple circulatory and physiological adaptations to enable them not to breathe for long periods.[50]
Smaller pond turtles, like these Northern red-bellied cooters, regulate their temperature by basking in the sun.
A green sea turtle grazing on seagrass
The oblong turtle has a sizable vocal repertoire.[65]
Crested caracara eating a turtle
Desert tortoises fighting
Mounting behavior in the three-toed box turtle
A female common snapping turtle depositing her eggs in a hole she dug
Marginated tortoise emerges from its egg
Diagram of the origins of the turtle body plan through the Triassic: isolated bony plates evolved to form a complete shell, in a sequence involving Pappochelys, Eorhynchochelys, Odontochelys, and Proganochelys.[18]
Neck retraction
Pleurodira retract their necks sideways.
The different mechanisms of neck retraction in the two suborders of turtles
Many turtles have been killed accidentally in fishing nets.[134] Some trawlers now use nets fitted with turtle excluders.[135] Seen here, a loggerhead escapes a net so fitted.