Laemophloeidae


Acompsophloeus Thomas, 2010
Blubos Lefkovitch, 1962
Brontolaemus Sharp, 1885
Carinophloeus Lefkovitch, 1961
Caulonomus Wollaston, 1862
Charaphloeus Casey, 1916
Cryptolestes Ganglbauer, 1899
Cucujinus Arrow, 1920
Deinophloeus Sharp, 1899
Dysmerus Casey, 1884
Gannes Lefkovitch, 1962
Heterojinus Sengupta & Mukhopadhyay, 1978
Laemophloeus Laporte de Castelnau, 1840
Lathropus Erichson, 1845
Lepidophloeus Thomas, 1984
Leptophloeus Casey, 1916
Magnoleptus Lefkovitch, 1962
Mariolaemus Lefkovitch, 1962
Mestolaemus Lefkovitch, 1962
Metaxyphloeus Thomas, 1984
Microbrontes Reitter, 1874
Microlaemus Lefkovich, 1962
Narthecius J.LeConte, 1861
Nipponophloeus Sasaji, 1983
Notolaemus Lefkovitch, 1959
Odontophloeus Thomas, 1984
Parandrita J.LeConte & Horn, 1880
Paraphloeolaemus Thomas, 2018
Passandrophloeus Kessel, 1921
Phloeipsius Casey, 1916
Phloeolaemus Casey, 1916
Placonotus MacLeay, 1871
Planolestes Lefkovitch, 1958
Pseudophloeus Yablokov-Khnzoryan, 1977
Rhabdophloeus Sharp, 1899
Rhinolaemus Steel, 1954
Rhinomalus Gemm, 1870
Rhinophloeus Sharp, 1899
Sinuatophloeus Kessel, 1921
Xylolestes Lefkovitch, 1962
Xylophloeus Lefkovitch, 1962

Laemophloeidae, "lined flat bark beetles," is a family in the superfamily Cucujoidea characterized by predominantly dorso-ventrally compressed bodies, head and pronotal discs bordered by ridges or grooves, and inverted male genitalia.[1][2] Size range of adults is 1–5 mm (0.04–0.2 in) in length. Currently, it contains 40 genera and about 450 species,[3] and is represented on all continents except Antarctica; species richness is greatest in the tropics.

Historically, Laemophloeidae was treated as a subfamily of Cucujidae, but starting in the middle of the 20th century,[4] most of what had been treated as subfamilies of the Cucujidae were considered to be families.

Most laemophloeids, adults and larvae, are found under bark of dead trees, where they apparently are primarily fungivores,[5] although some genera with adults having subcylindrical bodies (e.g., Leptophloeus, Dysmerus) occur in the galleries of bark beetles (Curculionidae: Scolytinae), upon which they may feed.[6] A few genera, but most particularly Cryptolestes, contain some species that are pests of stored grain products. The most important of these are Cryptolestes ferrugineus (Stephens), Cryptolestes pusillus (Schönherr), and Cryptolestes turcicus (Grouvelle).[7]

Several genera exhibit unusual modifications to male antennae (especially Cryptolestes, Dysmerus, and Microbrontes), with the scape expanded into hook-like or blade-like structures.[8][9] Several other genera (Rhinomalus, Rhinophloeus, and Metaxyphloeus) related to Laemophloeus are atypical in that the adults are rostrate to varying degrees.[10] Photographs of most world genera are available at,[11] and most North American species are pictured at:[12]

According to an article published by PLOS ONE, a discovery has been made regarding the larvae of laemophloeidae. Studies show that when the larvae is in the open, the larvae will straighten themselves out and jump curling up into a ball in the process. It has also been discovered that the larvae can bounce when they are curled up.[13][14]