Striped hyena


The striped hyena (Hyaena hyaena) is a species of hyena native to North and East Africa, the Middle East, the Caucasus, Central Asia and the Indian subcontinent. It is the only extant species in the genus Hyaena. It is listed by the IUCN as near-threatened, as the global population is estimated to be under 10,000 mature individuals which continues to experience deliberate and incidental persecution along with a decrease in its prey base such that it may come close to meeting a continuing decline of 10% over the next three generations.[1] It is also the national animal of Lebanon.[4]

It is the smallest of the true[clarification needed] hyenas and retains many primitive viverrid characteristics lost in larger species,[5] having a smaller and less specialised skull.[6][7] Though primarily a scavenger, large specimens have been known to kill their own prey,[8] and attacks on humans have occurred in rare instances.[9] The striped hyena is a monogamous animal, with both males and females assisting one another in raising their cubs.[10] A nocturnal animal, the striped hyena typically only emerges in complete darkness, and is quick to return to its lair before sunrise.[11] Although it has a habit of feigning death when attacked, it has been known to stand its ground against larger predators in disputes over food.[12]

The striped hyena features prominently in Middle Eastern and Asian folklore. In some areas, its body parts are considered magical, and are used as charms or talismans.[13] It is mentioned in the Hebrew Bible, where it is referred to as tzebua or zevoa, though it is absent in some Bible translations into English.[14] Ancient Greeks knew it as γλάνος (glános) and ύαινα (húaina) and were familiar with it from the Aegean coast of Asia Minor.[15]

The species may have evolved from H. namaquensis of Pliocene Africa. Striped hyena fossils are common in Africa, with records going back as far as the Middle Pleistocene and even to the Villafranchian. As fossil striped hyenas are absent from the Mediterranean region, it is likely that the species is a relatively late invader to Eurasia, having likely spread outside Africa only after the extirpation of spotted hyenas from Asia at the end of the last glacial period. The striped hyena occurred for some time in Europe during the Pleistocene, having been particularly widespread in France and Germany. It also occurred in Montmaurin, Hollabrunn in Austria, the Furninha Cave in Portugal and the Genista Caves in Gibraltar. The European form was similar in appearance to modern populations, but was larger, being comparable in size to the brown hyena.[5]


Skull drawn by V. N. Lyakhov.
Dentition, as illustrated in Knight's Sketches in Natural History.
Skeleton
A pair of striped hyenas fighting at Colchester Zoo.
Illustration from Frank Finn's Wild Beasts of the World (1909).
Stuffed striped hyena defending a sheep carcass from hooded crows, exhibited in The Museum of Zoology, St. Petersburg.
Illustration of striped hyena (top) and spotted hyena (bottom).
A wild individual at Blackbuck National Park, Velavadar, Gujarat state, India.
Striped hyena pugmark in wet clay, Coimbatore, Tamil Nadu, India.
A striped hyena, as depicted on the Nile mosaic of Palestrina.
Striped hyena scavenging on poultry waste in Dahod district, Gujarat, India.
Engraving of a striped hyena attacking a man in The Naturalist's Cabinet (1806).
Hyena (1739) by Jean-Baptiste Oudry.
A striped hyena being speared in British India, as illustrated in the Illustrated London News.
A tame striped hyena.