Elapidae


Elapidae (/ɪˈlæpɪd/, commonly known as elapids /ˈɛləpɪdz/; Ancient Greek: ἔλλοψ éllops "sea-fish"[6]) is a family of venomous snakes characterized by their permanently erect fangs at the front of the mouth. Many members of this family are also recognized by their threat display of rearing upwards while spreading the neck-flap. Elapids are endemic to tropical and subtropical regions around the world, with terrestrial forms in Asia, Australia, Africa, and the Americas and marine forms in the Pacific and Indian Oceans. Members of the family have a wide range of sizes, from the 18 cm (7.1 in) white-lipped snake to the 5.85 m (19 ft 2 in) king cobra. Most species have neurotoxins in their venom which is channeled by their hollow fangs, while some may contain other toxic components in various proportions. The family includes 55 genera with some 360 species and over 170 subspecies.

Terrestrial elapids look similar to the Colubridae; almost all have long, slender bodies with smooth scales, a head covered with large shields and not always distinct from the neck, and eyes with round pupils. In addition, their behavior is usually quite active, and most are oviparous. Exceptions to all these generalizations occur: e.g. the death adders (Acanthophis) include short and fat, rough-scaled, very broad-headed, cat-eyed, live-bearing, sluggish ambush predators with partly fragmented head shields.

Sea snakes (the Hydrophiinae), sometimes considered to be a separate family, have adapted to a marine way of life in different ways and to various degrees. All have evolved paddle-like tails for swimming and the ability to excrete salt. Most also have laterally compressed bodies, their ventral scales are much reduced in size, their nostrils are located dorsally (no internasal scales), and they give birth to live young (viviparity). The reduction in ventral scaling has greatly diminished their terrestrial mobility, but aids in swimming.

Members of this family have a wide range of sizes. Drysdalia species are small serpents typically 50 cm (20 in) and down to 18 cm (7.1 in) in length. Cobras, mambas, and taipans are mid- to large sized snakes which can reach 2 m (6 ft 7 in) or above. The king cobra is the world's longest venomous snake with a maximum length of 5.85 m (19.2 ft) and an average mass of 6 kg (13 lb).[7]

All elapids have a pair of proteroglyphous fangs to inject venom from glands located towards the rear of the upper jaw. The fangs, which are enlarged and hollow, are the first two teeth on each maxillary bone. Usually only one fang is in place on each side at any time. The maxilla is intermediate in both length and mobility between typical colubrids (long, less mobile) and viperids (very short, highly mobile). When the mouth is closed, the fangs fit into grooved slots in the buccal floor and usually below the front edge of the eye and are angled backwards; some elapids (Acanthophis, taipan, mamba, and king cobra) have long fangs on quite mobile maxillae and can make fast strikes. A few species are capable of spraying their venom from forward-facing holes in their fangs for defense, as exemplified by spitting cobras.

Most elapids are terrestrial, while some are strongly arboreal (African Pseudohaje and Dendroaspis, Australian Hoplocephalus). Many species are more or less specialized burrowers (e.g. Ogmodon, Parapistocalamus, Simoselaps, Toxicocalamus, and Vermicella) in either humid or arid environments. Some species have very generalised diets (euryphagy), but many taxa have narrow prey preferences (stenophagy) and correlated morphological specializations, for example feeding almost exclusively on other serpents (especially the king cobra and kraits). Elapids may display a series of warning signs if provoked, either obviously or subtly. Cobras and mambas lift their inferior body parts, expand hoods, and hiss if threatened; kraits often curl up before hiding their heads down their bodies.


The lateral view of a king cobra's skull showing fangs