Baryonyx


Baryonyx (/ˌbæriˈɒnɪks/) is a genus of theropod dinosaur which lived in the Barremian stage of the Early Cretaceous period, about 130–125 million years ago. The first skeleton was discovered in 1983 in the Smokejack Clay Pit, of Surrey, England, in sediments of the Weald Clay Formation, and became the holotype specimen of Baryonyx walkeri, named by palaeontologists Alan J. Charig and Angela C. Milner in 1986. The generic name, Baryonyx, means "heavy claw" and alludes to the animal's very large claw on the first finger; the specific name, walkeri, refers to its discoverer, amateur fossil collector William J. Walker. The holotype specimen is one of the most complete theropod skeletons from the UK (and remains the most complete spinosaurid), and its discovery attracted media attention. Specimens later discovered in other parts of the United Kingdom and Iberia have also been assigned to the genus, though many have since been moved to new genera.

The holotype specimen, which may not have been fully grown, was estimated to have been between 7.5 and 10 metres (25 and 33 feet) long and to have weighed between 1.2 and 1.7 metric tons (1.3 and 1.9 short tons; 1.2 and 1.7 long tons). Baryonyx had a long, low, and narrow snout, which has been compared to that of a gharial. The tip of the snout expanded to the sides in the shape of a rosette. Behind this, the upper jaw had a notch which fitted into the lower jaw (which curved upwards in the same area). It had a triangular crest on the top of its nasal bones. Baryonyx had a large number of finely serrated, conical teeth, with the largest teeth in front. The neck formed an S-shape, and the neural spines of its dorsal vertebrae increased in height from front to back. One elongated neural spine indicates it may have had a hump or ridge along the centre of its back. It had robust forelimbs, with the eponymous first-finger claw measuring about 31 centimetres (12 inches) long.

Now recognised as a member of the family Spinosauridae, Baryonyx's affinities were obscure when it was discovered. Some researchers have suggested that Suchosaurus cultridens is a senior synonym (being an older name), and that Suchomimus tenerensis belongs in the same genus; subsequent authors have kept them separate. Baryonyx was the first theropod dinosaur demonstrated to have been piscivorous (fish-eating), as evidenced by fish scales in the stomach region of the holotype specimen. It may also have been an active predator of larger prey and a scavenger, since it also contained bones of a juvenile iguanodontid. The creature would have caught and processed its prey primarily with its forelimbs and large claws. Baryonyx may have had semiaquatic habits, and coexisted with other theropod, ornithopod, and sauropod dinosaurs, as well as pterosaurs, crocodiles, turtles and fishes, in a fluvial environment.


Replica of a large, thick claw
Cast of the hand claw that the name Baryonyx was based on, in Palais de la Découverte, Paris
Skeletal elements of the holotype specimen as archived (above), and reconstructed skeleton on exhibit (below), in Natural History Museum, London
Skeletal diagram of Baryonyx with the bones of Portuguese specimen ML1190 shown in red; it was originally considered a specimen of Baryonyx, but was since moved to Iberospinus
1878 lithograph (left) and modern photo showing the holotype tooth of Suchosaurus cultridens, which may represent the same animal as B. walkeri
Size of various spinosaurids (Baryonyx in yellow, second from left) compared with a human
Snout of the holotype specimen, from the left and below
Holotype dentary from the left
Reconstruction of the holotype skull, Museon, The Hague
Life restoration
Three cervical vertebrae from the neck of the holotype in left side view, the third also shown from the front (left), and reconstructed necks of the spinosaurids Sigilmassasaurus (A) and Baryonyx (B), showing their curvature (right)
Snouts of Cristatusaurus (A-C), Suchomimus (D-I), and Baryonyx (J-L)
Reconstructed skeleton in Japan
Skeletal diagram of the holotype specimen (below) compared with the related genus Suchomimus
Distribution of spinosaurids in Europe and North Africa during the Cretaceous; 1, 3, 4, 5, 6 are Baryonyx
Restoration of Baryonyx with a fish
Abraded bones of a young iguanodontid found within the rib cage of the B. walkeri holotype
Video showing a CT scan 3D model of the holotype snout in rotation (left), and three skulls of extant crocodilians that were compared to those of spinosaurs in a 2013 study (right)
Resting Baryonyx being groomed by small pterosaurs and birds
Spatial distribution of abelisaurids, carcharodontosaurids, and spinosaurids (the latter strongly associated with coastal environments)
Restoration of Baryonyx with a fish
Diagram showing known material of Riparovenator (front) and Ceratosuchops (rear), spinosaurids from the Wessex Formation which may have lived alongside Baryonyx
Model carcass based on the position of the holotype bones, NHM