Passerine


A passerine (/ˈpæsərn/) is any bird of the order Passeriformes (/ˈpæsərɪfɔːrmz/; from Latin passer 'sparrow' and formis '-shaped'), which includes more than half of all bird species. Sometimes known as perching birds or songbirds, passerines are distinguished from other orders of birds by the arrangement of their toes (three pointing forward and one back), which facilitates perching.

With more than 140 families and some 6,500 identified species,[1] Passeriformes is the largest order of birds and among the most diverse orders of terrestrial vertebrates, representing 60% of birds.[2][3] Passerines are divided into three suborders: Acanthisitti (New Zealand wrens), Tyranni (suboscines), and Passeri (oscines).[4][5] The passerines contain several groups of brood parasites such as the viduas, cuckoo-finches, and the cowbirds. Most passerines are omnivorous, while the shrikes are carnivorous.

The terms "passerine" and "Passeriformes" are derived from the scientific name of the house sparrow, Passer domesticus, and ultimately from the Latin term passer, which refers to sparrows and similar small birds.

The order is divided into three suborders, Tyranni (suboscines), Passeri (oscines), and the basal Acanthisitti.[6] Oscines have the best control of their syrinx muscles among birds, producing a wide range of songs and other vocalizations, though some of them, such as the crows, do not sound musical to human beings. Some, such as the lyrebird, are accomplished imitators.[7] The New Zealand wrens are tiny birds restricted to New Zealand, at least in modern times; they were long placed in Passeri.

Most passerines are smaller than typical members of other avian orders. The heaviest and altogether largest passerines are the thick-billed raven[8] and the larger races of common raven, each exceeding 1.5 kg (3.3 lb) and 70 cm (28 in). The superb lyrebird and some birds-of-paradise, due to very long tails or tail coverts, are longer overall. The smallest passerine is the short-tailed pygmy tyrant, at 6.5 cm (2.6 in) and 4.2 g (0.15 oz).[9]

The foot of a passerine has three toes directed forward and one toe directed backward, called anisodactyl arrangement, and the hind toe (hallux) joins the leg at approximately the same level as the front toes. This arrangement enables passerine birds to easily perch upright on branches. The toes have no webbing or joining, but in some cotingas, the second and third toes are united at their basal third.


Pterylosis or the feather tracts in a typical passerine
Male superb lyrebird (Menura novaehollandiae): This very primitive songbird shows strong sexual dimorphism, with a peculiarly apomorphic display of plumage in males.
Wieslochia fossil
New Zealand rock wren (Xenicus gilviventris), one of the two surviving species of suborder Acanthisitti
Javan banded pitta (Hydrornis guajanus), an Old World suboscine.
Andean cock-of-the-rock (Rupicola peruvianus) a New World suboscine
Male stitchbird or hihi (Notiomystis cincta) showing convergence with honeyeaters
Male regent bowerbird (Sericulus chrysocephalus, Ptilonorhynchidae)
Tiny goldcrest (Regulus regulus) belongs to a minor but highly distinct lineage of Passeri
Reed warblers, such as this Blyth's reed warbler (Acrocephalus dumetorum), are now in the Acrocephalidae
Eurasian blue tit (Cyanistes caeruleus) and its relatives stand well apart from rest of the Sylvioidea sensu lato
Brown-headed nuthatch (Sitta pusilla), nuthatches can climb downwards head-first
Hermit thrush (Catharus guttatus), like many Muscicapoidea a stout and cryptic bird with complex vocalizations.
Like these male (right) and female Gouldian finches (Erythrura gouldiae), many Passeroidea are very colorful
Lesser striped swallow (Cecropis abyssinica), showing some apomorphies of its ancient yet highly advanced lineage.
Gran Canaria blue chaffinch (male)