Eolambia


Eolambia (meaning "dawn lambeosaurine") is a genus of herbivorous hadrosauroid dinosaur from the early Late Cretaceous of the United States. It contains a single species, E. caroljonesa, named by paleontologist James Kirkland in 1998. The type specimen of Eolambia was discovered by Carole and Ramal Jones in 1993; the species name honors Carole. Since then, hundreds of bones have been discovered from both adults and juveniles, representing nearly every element of the skeleton. All of the specimens have thus far been found in Emery County, Utah, in a layer of rock known as the Mussentuchit Member of the Cedar Mountain Formation.

Measuring up to 6 meters (20 ft) long, Eolambia is a large member of its group. While it closely approaches the Asian hadrosauroids Equijubus, Probactrosaurus, and Choyrodon, in traits of the skull, vertebrae, and limbs, it may actually be more closely related to the North American Protohadros. This grouping, based on the straightness of the quadrate bone and scapula, would represent an isolated, endemic radiation of hadrosauroids. Despite resembling hadrosaurids – lambeosaurine hadrosaurids in particular – in several features, leading to its initial identification as one of them, these similarities have been rejected as either entirely convergent or misinterpreted.

Eolambia would have lived in a forested environment at the edge of lakes in a humid floodplain environment, feeding on gymnosperms, ferns, and flowering plants. The water levels in the lakes changed over time with cyclical wet and dry spells caused by the precession of the Earth, reflected by alternating bands in the sediments of the Mussentuchit Member. As a juvenile, Eolambia would have been preyed upon by large crocodylomorphs residing in the lake waters. With increasing age, however, they became impervious to the crocodylomorphs, and mature individuals (at least eight to nine years in age) were preyed on by large theropods such as the neovenatorid Siats.

In 1979 Peter Galton and James A. Jensen described a fragmentary right femur, BYU 2000, belonging to a hadrosaurian dinosaur discovered in sediments belonging to the Cedar Mountain Formation in Arches National Park, Utah. Though poor material, it was important for it (alongside a second North American femur described in the paper) was the first hadrosaur specimen from the Lower Cretaceous anywhere in the world. Galton and Jensen hypothesized more complete remains of a hadrosaur may be found from the formation in the future.[1] Various hadrosauroid teeth had also been found in quarries of small vertebrates in the western region of the San Rafael Swell, near Castle Dale in Emery County, Utah; they were described in 1991 by J. Michael Parrish. Subsequently, in 1993, Carole Jones and her husband Ramal Jones discovered fragmentary bones in a fossil site located in the northwestern region of the Swell. They brought the site to the attention of Donald Burge, director of the institution then called the College of Eastern Utah Prehistoric Museum (CEUM). The site, which is formally known as CEUM Locality 42em366v, would subsequently be named Carol's Site (sic) in her honour. The fossils, stored under the specimen number CEUM 9758, represent the partial remains of an adult hadrosauroid, including parts of the skull, vertebrae, ischium, and leg. CEUM 5212, a partial skull and forelimb from an adult, was found nearby in CEUM Locality 42em369v.[2] CEUM 8786, a left femur from an adult, was discovered later in Carol's Site, and was not described until 2012.[3]


The dentary of the holotype Eolambia specimen CEUM 9758
Eolambia was named after Lambeosaurus due to its initial interpretation as a basal lambeosaurine
Specimen CEUM 34447, an unusually deep dentary which may or may not pertain to Eolambia
Specimen FMNH PR 3847, discovered by the Field Museum of Natural History in 2008
The sandstone block containing specimen FMNH PR 3847; these rocks are typical of the Mussentuchit Member
A restoration of Eolambia caroljonesa based on skeletal diagrams and related species
Maxillae of Eolambia
Dentary of Eolambia
Cervical vertebrae of Eolambia, showing unusual third cervical (A)
Dorsal vertebrae of Eolambia, showing transition in location of vertebral processes
Shoulder girdle elements of Eolambia, including scapula (C-D)
Lower leg of Eolambia
Teeth of Eolambia
Caudal vertebrae of Eolambia; note the tall neural spines
Pelvic girdle elements of Eolambia – ilium (A-B), ischium (C-D), pubis (E-F)
Sacrum of Eolambia; note the groove on the bottom surface (A)
Premaxilla of Eolambia
Skull of Probactrosaurus, the closest relative of Eolambia according to Head
Skull of Altirhinus, considered by Norman as the sister group of Eolambia but now known to be more basal
Quadrate of Eolambia
Reconstruction of Jintasaurus, recovered by Zheng et al. as a sister group of Eolambia
Cross section of a rib from specimen FMNH PR 3847; LAGs marked by arrows
Skeletal diagram comparing the known material of the holotype adult (right) and a juvenile individual (left)
The precession of the Earth influenced the local climate
Shoreline debris at the present-day Rathtrevor Beach Provincial Park
Pholidosaurids like Sarcosuchus, shown here, might have lived alongside Eolambia
Skeleton of the co-existing nodosaurid Animantarx
Leg bones of Eolambia – femur (A-F), tibia (G-L), fibula (M-N)