Ichthyosaur


Ichthyosaurs (Ancient Greek for "fish lizard" – ἰχθύς or ichthys meaning "fish" and σαῦρος or sauros meaning "lizard") are large extinct marine reptiles. Ichthyosaurs belong to the order known as Ichthyosauria or Ichthyopterygia ('fish flippers' – a designation introduced by Sir Richard Owen in 1842, although the term is now used more for the parent clade of the Ichthyosauria).

Ichthyosaurs thrived during much of the Mesozoic era; based on fossil evidence, they first appeared around 250 million years ago (Ma) and at least one species survived until about 90 million years ago,[1][2] into the Late Cretaceous. During the Early Triassic epoch, ichthyosaurs evolved from a group of unidentified land reptiles that returned to the sea, in a development similar to how the mammalian land-dwelling ancestors of modern-day dolphins and whales returned to the sea millions of years later, which they gradually came to resemble in a case of convergent evolution. Ichthyosaurs were particularly abundant in the Late Triassic and Early Jurassic periods, until they were replaced as the top aquatic predators by another marine reptilian group, the Plesiosauria, in the later Jurassic and Early Cretaceous, though previous views of ichthyosaur decline during this period are probably overstated. Ichthyosaurs diversity declined due to environmental volatility caused by climatic upheavals in the early Late Cretaceous, becoming extinct around the Cenomanian-Turonian boundary.

Science became aware of the existence of ichthyosaurs during the early nineteenth century, when the first complete skeletons were found in England. In 1834, the order Ichthyosauria was named. Later that century, many excellently preserved ichthyosaur fossils were discovered in Germany, including soft-tissue remains. Since the late twentieth century, there has been a revived interest in the group, leading to an increased number of named ichthyosaurs from all continents, with over fifty valid genera being now known.

Ichthyosaur species varied from 1 to 20 metres (3 to 66 ft) in length. Ichthyosaurs resembled both modern fish and dolphins. Their limbs had been fully transformed into flippers, which sometimes contained a very large number of digits and phalanges. At least some species possessed a dorsal fin. Their heads were pointed, and the jaws often were equipped with conical teeth that could help to catch smaller prey. Some species had larger, bladed teeth with which they could attack large animals. The eyes were very large, probably useful when deep diving. The neck was short, and later species had a rather stiff trunk. These also had a more vertical tail fin, used for a powerful propulsive stroke. The vertebral column, made of simplified disc-like vertebrae, continued into the lower lobe of the tail fin. Ichthyosaurs were air-breathing, warm-blooded,and bore live young. They may have had a layer ofblubber for insulation.


Ichthyosaur and Plesiosaur by Édouard Riou, 1863. (This old representation of a plesiosaur lifting its head is not accurate and physically impossible)
The skull found by Joseph Anning in 1811[8]
The torso found by Mary Anning in 1812[8]
drawing of side view of the skeleton of an animal with a long thin skull, long tail, and paddles
Diagram of the skeletal anatomy of Ichthyosaurus communis from an 1824 paper by Conybeare
"Professor Ichthyosaurus" shows his pupils the skull of extinct man, caricature of Charles Lyell by Henry De la Beche (1830)
Hawkins' specimens are still the showpieces of the Natural History Museum
Ichthyosaurus figure at Crystal Palace Park (1854)
Typical Holzmaden fossils: adult and juvenile Stenopterygius quadriscissus
Precious opal replacing ichthyosaur backbone, display specimen, South Australian Museum
Grippia longirostris from the early Triassic of Spitsbergen was already well-adapted to an aquatic lifestyle.
Hupehsuchus
Mixosaurus
Shonisaurus popularis
Stenopterygius resembled a modern dolphin.
Many ichthyosaur lineages continued into the Cretaceous.
Restoration of Platypterygius kiprijanovi – Albian-Cenomanian of Kursk region (Russia)
CGI restoration of Ichthyosaurus communis
Examples of distinct features shared both by dolphins and derived ichthyopterygians
The skull of Temnodontosaurus platyodon has the typical ichthyosaurian shape with an elongated snout and large eye sockets.
Ichthyosaur vertebra from the Sundance Formation (Jurassic) of Natrona County, Wyoming: Note the characteristic hourglass cross-section. (Scale in mm.)
In this specimen seen from below, what looks like a breastbone is in fact the fused coracoids
Ichthyosaur 'paddle' (Charmouth Heritage Coast Centre)
In this arm of Ophthalmosaurus icenius, an additional upper row of elements has developed, ending above in an extra lower arm bone.
A Holzmaden ichthyosaur in which the preparer found organic remains in the position of the dorsal fin, but failed to locate any for the flippers.
An ichthyosaur coprolith
Temnodontosaurus acutirostris with ammonoids
Caypullisaurus is attacked by the crocodylomorph Dakosaurus
Despite their considerable size, the flippers of the Amazon river dolphin are mainly used as rudders.
Temnodontosaurus had the largest eyes of any known vertebrate, indicating a good diving capacity
Chaohusaurus with three juveniles
Detail of a female Stenopterygius with a great number of fetuses in her belly, one of which has been expelled