Bat


Bats are mammals of the order Chiroptera.[a] With their forelimbs adapted as wings, they are the only mammals capable of true and sustained flight. Bats are more maneuverable than most birds, flying with their very long spread-out digits covered with a thin membrane or patagium. The smallest bat, and arguably the smallest extant mammal, is Kitti's hog-nosed bat, which is 29–34 millimetres (1+181+38 inches) in length, 150 mm (6 in) across the wings and 2–2.6 g (116332 oz) in mass. The largest bats are the flying foxes, with the giant golden-crowned flying fox, Acerodon jubatus, reaching a weight of 1.6 kg (3+12 lb) and having a wingspan of 1.7 m (5 ft 7 in).

The second largest order of mammals after rodents, bats comprise about 20% of all classified mammal species worldwide, with over 1,400 species. These were traditionally divided into two suborders: the largely fruit-eating megabats, and the echolocating microbats. But more recent evidence has supported dividing the order into Yinpterochiroptera and Yangochiroptera, with megabats as members of the former along with several species of microbats. Many bats are insectivores, and most of the rest are frugivores (fruit-eaters) or nectarivores (nectar-eaters). A few species feed on animals other than insects; for example, the vampire bats feed on blood. Most bats are nocturnal, and many roost in caves or other refuges; it is uncertain whether bats have these behaviours to escape predators. Bats are present throughout the world, with the exception of extremely cold regions. They are important in their ecosystems for pollinating flowers and dispersing seeds; many tropical plants depend entirely on bats for these services.

Bats provide humans with some direct benefits, at the cost of some disadvantages. Bat dung has been mined as guano from caves and used as fertiliser. Bats consume insect pests, reducing the need for pesticides and other insect management measures. They are sometimes numerous enough and close enough to human settlements to serve as tourist attractions, and they are used as food across Asia and the Pacific Rim. However, fruit bats are frequently considered pests by fruit growers. Due to their physiology, bats are one type of animal that acts as a natural reservoir of many pathogens, such as rabies; and since they are highly mobile, social, and long-lived, they can readily spread disease among themselves. If humans interact with bats, these traits become potentially dangerous to humans.

Depending on the culture, bats may be symbolically associated with positive traits, such as protection from certain diseases or risks, rebirth, or long life, but in the West, bats are popularly associated with darkness, malevolence, witchcraft, vampires, and death.

An older English name for bats is flittermouse, which matches their name in other Germanic languages (for example German Fledermaus and Swedish fladdermus), related to the fluttering of wings. Middle English had bakke, most likely cognate with Old Swedish natbakka ("night-bat"), which may have undergone a shift from -k- to -t- (to Modern English bat) influenced by Latin blatta, "moth, nocturnal insect". The word "bat" was probably first used in the early 1570s.[2][3] The name "Chiroptera" derives from Ancient Greek: χείρ – cheir, "hand"[4] and πτερόν – pteron, "wing".[1][5]

The delicate skeletons of bats do not fossilise well; it is estimated that only 12% of bat genera that lived have been found in the fossil record.[6] Most of the oldest known bat fossils were already very similar to modern microbats, such as Archaeopteropus (32 million years ago).[7] The extinct bats Palaeochiropteryx tupaiodon (48 million years ago) and Hassianycteris kumari (48 million years ago) are the first fossil mammals whose colouration has been discovered: both were reddish-brown.[8][9]


The early Eocene fossil microchiropteran Icaronycteris, from the Green River Formation
Giant golden-crowned flying fox, Acerodon jubatus
"Chiroptera" from Ernst Haeckel's Kunstformen der Natur, 1904
A preserved megabat showing how the skeleton fits inside its skin
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Little brown bat take off and flight
Wing membranes (patagia) of Townsend's big-eared bat, Corynorhinus townsendii
Group of megabats roosting
The wings are highly vascularized membranes, the larger blood vessels visible against the light.[68]
Principle of bat echolocation: orange is the call and green is the echo.
The tiger moth (Bertholdia trigona) can jam bat echolocation.[88][89]
Thermographic image of a bat using trapped air as insulation
A tricoloured bat (Perimyotis subflavus) in torpor
Tent-making bats (Uroderma bilobatum) in Costa Rica
Mexican long-tongued bat (Choeronycteris mexicana) drinking from an agave flower
An Egyptian fruit bat (Rousettus aegyptiacus) carrying a fig
The greater noctule bat (Nyctalus lasiopterus) uses its large teeth to catch birds.[156]
The common vampire bat (Desmodus rotundus) feeds on blood (hematophagy).
A little brown bat with white nose syndrome
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Bracken Bat Cave, home to twenty million Mexican free-tailed bats
Acoustics of the songs of Mexican free-tailed bats[210]
Group of polygynous vampire bats
Newborn common pipistrelle, Pipistrellus pipistrellus
The bat scientist Lauri Lutsar is checking the age of the bat he is holding as part of a national monitoring program in Estonia

Conservation statuses of bats as of 2020 according to the IUCN (1,314 species in total)[235]

  Critically endangered (1.6%)
  Endangered (6.3%)
  Vulnerable (8.3%)
  Near-threatened (6.7%)
  Least concern (58.0%)
  Data deficient (18.4%)
  Extinct (0.7%)
Francisco Goya, The Sleep of Reason Produces Monsters, 1797
Zapotec bat god, Oaxaca, 350–500 CE