Amphibian


Amphibians are ectothermic, tetrapod vertebrates of the class Amphibia. All living amphibians belong to the group Lissamphibia. They inhabit a wide variety of habitats, with most species living within terrestrial, fossorial, arboreal or freshwater aquatic ecosystems. Thus amphibians typically start out as larvae living in water, but some species have developed behavioural adaptations to bypass this.

The young generally undergo metamorphosis from larva with gills to an adult air-breathing form with lungs. Amphibians use their skin as a secondary respiratory surface and some small terrestrial salamanders and frogs lack lungs and rely entirely on their skin. They are superficially similar to lizards but, along with mammals and birds, reptiles are amniotes and do not require water bodies in which to breed. With their complex reproductive needs and permeable skins, amphibians are often ecological indicators; in recent decades there has been a dramatic decline in amphibian populations for many species around the globe.

The earliest amphibians evolved in the Devonian period from sarcopterygian fish with lungs and bony-limbed fins, features that were helpful in adapting to dry land. They diversified and became dominant during the Carboniferous and Permian periods, but were later displaced by reptiles and other vertebrates. Over time, amphibians shrank in size and decreased in diversity, leaving only the modern subclass Lissamphibia.

The three modern orders of amphibians are Anura (the frogs), Urodela (the salamanders), and Apoda (the caecilians). The number of known amphibian species is approximately 8,000, of which nearly 90% are frogs. The smallest amphibian (and vertebrate) in the world is a frog from New Guinea (Paedophryne amauensis) with a length of just 7.7 mm (0.30 in). The largest living amphibian is the 1.8 m (5 ft 11 in) South China giant salamander (Andrias sligoi), but this is dwarfed by the extinct 9 m (30 ft) Prionosuchus from the middle Permian of Brazil. The study of amphibians is called batrachology, while the study of both reptiles and amphibians is called herpetology.

The word amphibian is derived from the Ancient Greek term ἀμφίβιος (amphíbios), which means 'both kinds of life', ἀμφί meaning 'of both kinds' and βιος meaning 'life'. The term was initially used as a general adjective for animals that could live on land or in water, including seals and otters.[3] Traditionally, the class Amphibia includes all tetrapod vertebrates that are not amniotes. Amphibia in its widest sense (sensu lato) was divided into three subclasses, two of which are extinct:[4]

The actual number of species in each group depends on the taxonomic classification followed. The two most common systems are the classification adopted by the website AmphibiaWeb, University of California, Berkeley and the classification by herpetologist Darrel Frost and the American Museum of Natural History, available as the online reference database "Amphibian Species of the World".[6] The numbers of species cited above follows Frost and the total number of known amphibian species as of March 31, 2019 is exactly 8,000,[7] of which nearly 90% are frogs.[8]


Many amphibians — like this Ceratophrys cranwelli — exhibit biofluorescence.[2]
The world's smallest known vertebrate, Paedophryne amauensis, sitting on a U.S. dime. The dime is 17.9 mm in diameter, for scale
Triadobatrachus massinoti, a proto-frog from the Early Triassic of Madagascar
Top: Restoration of Eusthenopteron, a fully aquatic lobe-finned fishBottom: Restoration of Tiktaalik, an advanced tetrapodomorph fish
The Permian lepospondyl Diplocaulus was largely aquatic
The temnospondyl Eryops had sturdy limbs to support its body on land
Red-eyed tree frog (Agalychnis callidryas) with limbs and feet specialised for climbing
Japanese giant salamander
(Andrias japonicus), a primitive salamander
Danube crested newt
(Triturus dobrogicus), an advanced salamander
The limbless South American caecilian Siphonops paulensis
The bright colours of the common reed frog (Hyperolius viridiflavus) are typical of a toxic species
Cross section of frog skin.
  1. Mucous gland
  2. Chromatophore
  3. Granular poison gland
  4. Connective tissue
  5. Stratum corneum
  6. Transition zone
  7. Epidermis
  8. Dermis
Skull diagram of Xenotosuchus, a temnospondyl
Skeleton of the Surinam horned frog
(Ceratophrys cornuta)
Skull of Yaksha perettii an extinct albanerpetontid
Didactic model of an amphibian heart.
Juvenile amphibian circulatory systems are single loop systems which resemble fish.
  1. Internal gills where the blood is reoxygenated
  2. Point where the blood is depleted of oxygen and returns to the heart via veins
  3. Two chambered heart
Red indicates oxygenated blood, and blue represents oxygen depleted blood.
Dissected frog:
  1. Right atrium
  2. Liver
  3. Aorta
  4. Egg mass
  5. Colon
  6. Left atrium
  7. Ventricle
  8. Stomach
  9. Left lung
  10. Spleen
  11. Small intestine
  12. Cloaca
The axolotl (Ambystoma mexicanum) retains its larval form with gills into adulthood
Male orange-thighed frog (Litoria xanthomera) grasping the female during amplexus
Sexual selection has been studied in the red back salamander
Frogspawn, a mass of eggs surrounded by jelly
Amphibian egg:
  1. Jelly capsule
  2. Vitelline membrane
  3. Perivitelline fluid
  4. Yolk plug
  5. Embryo
Early stages in the development of the embryos of the common frog (Rana temporaria)
Successive stages in the development of common toad (Bufo bufo) tadpoles, finishing with metamorphosis
Larva of the long-toed salamander
(Ambystoma macrodactylum)
Larva of the long-toed salamander
(Ambystoma macrodactylum)
Larvae of the alpine newt
(Ichthyosaura alpestris)
The caecilian Ichthyophis glutinosus with eggs and developing embryo
The ringed caecilian (Siphonops annulatus) resembles an earthworm
Male common rocket frog (Colostethus panamensis) carrying tadpoles on his back
Male common midwife toad (Alytes obstetricans) carrying eggs
Northwestern salamander
(Ambystoma gracile) eating a worm
Edible frog (Pelophylax esculentus) exhibiting cannibalism
Audio showing Brazilian torrent frog males executing advertisement, peep, and squeal calls.
Male treefrog (Dendropsophus microcephalus) inflating his air sac as he calls
American toad, (Anaxyrus americanus) singing
The red back salamander (Plethodon cinereus) defends a territory against intruders.
Cane toad (Rhinella marina) with poison glands behind the eyes
The fire salamander (Salamandra salamandra), a toxic species, wears warning colours.
Perhaps the most poisonous animal in the world, the golden poison frog (Phyllobates terribilis) is endemic to Colombia.[145]
The extinct golden toad (Bufo periglenes), last seen in 1989
The Hula painted frog (Discoglossus nigriventer) was believed to be extinct but was rediscovered in 2011.