Arbutus
Arbutus is a genus of 12 accepted species[2] of flowering plants in the family Ericaceae,[3] native to warm temperate regions of the Mediterranean, western Europe, the Canary Islands and North America. The name Arbutus was taken from Latin, where it referred to Arbutus unedo.[4]
Arbutus are small trees or shrubs with red flaking bark and edible red berries.[5] Fruit development is delayed for about five months after pollination, so that flowers appear while the previous year's fruit are ripening.[5] Peak flowering for the genus is in April with peak fruiting in October.[6]
The smooth wood of the tree is mentioned by Theophrastus in his Enquiry into Plants (Historia Plantarum) as formerly being used to make weaving spindles. An article on Arbutus tree cultivation in al-Andalus is brought down in Ibn al-'Awwam's 12th-century agricultural work, Book on Agriculture.[7]
Members of the genus are called madrones or madronas in the United States, from the Spanish name madroño (strawberry tree). On the south coast of British Columbia, Canada, where the species is common, arbutus is commonly used or, rarely and locally, "tick tree".[8][9] All refer to the same species, Arbutus menziesii, native to the Pacific Northwest and Northern and Central California regions. It is Canada's only native broadleaved evergreen tree. Some species in the genera Epigaea, Arctostaphylos and Gaultheria were formerly classified in Arbutus. As a result of its past classification, Epigaea repens (mayflower) has an alternative common name of "trailing arbutus".
A study published in 2001 which analyzed ribosomal DNA from Arbutus and related genera suggests that Arbutus is paraphyletic and the Mediterranean Basin species of Arbutus are more closely related to Arctostaphylos, Arctous, Comarostaphylis, Ornithostaphylos and Xylococcus than to the western North American species of Arbutus, and that the split between the two groups of species occurred at the Paleogene/Neogene boundary.[10] The 12 species are as follows:[2]
Arbutus species are used as food plants by some Lepidoptera species including emperor moth, Pavonia pavonia and the madrone butterfly.[15] The distribution of the latter species is in fact heavily affected by the distribution of the madrone.[15] For Athenaios, it is the tree which Asclepiades of Myrlea talks about (Deiphnosophists, II.35)
